Eye Color
If you want to pass eye colors onto your cubs, I recommend using Opal Saltlicks to guarantee all the cubs in the litter having one specific eye color. They can be purchased in Monkey Business for 500

NCL Exclusive
These eye colors can only be found on NCLs in Explore.
AlbinoidCeladonCeramicChamotteCordovanGlassGlazeMosaic
SmaltiTerracottaTessera
Breed Unique
These eyes can only be result from 'fails' when breeding lions with specific NCL Exclusive eye colors. They act similarly to Primal Fangs failing from Primal. The eyes required to breed them are listed under the eye color in bold!
BurnishGlazeMireTerracottaFaienceCeramicDuntTesseraRose EbonySmaltiClayChamotte/TerracottaRose QuartzCordovan/Glaze
Combo Eyes
These eyes can be obtained by breeding certain factors to other factors or passed down through breeding. There is currently only one Combo Eye on Lioden. Factors are listed below in bold!
CosmicGalaxy or Jasper x Dawn, Draconid, Starshine, Stellar, or Sunglow
Sectoral Eyes
These eyes can be obtained by breeding the colors in their name together, or passed down through breeding. For example, for Sectoral Brown and Blue, you would breed a lioness with Blue eyes to a stud with Brown.
Sectoral Amber & IceSectoral Ice & GreenSectoral Red & JetSectoral Brown & BlueSectoral Green & BlueSectoral Yellow & GreenSectoral Pink & VioletSectoral Crystal & LavenderSectoral Tyrian & IndigoSectoral Sapphire & Albinoid

Eye Color Continued
This contains all the event eye colors! These are still breed-only, I just had to separate them for character limit reasons. Sorry!

February
These eyes can only be found on NCLs during the February Event after Tier 3 of the Flirt Shop has been unlocked and passed down through breeding.
DoveMaroonPearlRose
April
These eyes can only be found on NCLs during the April Event after Tier 2 of the Egg Den has been unlocked and passed down through breeding.
DaisyGazaniaHyacinthIris

May
These eyes can only be found on May Groupies during the May Event and passed down through breeding. Hetero eyes have the 'Hetero' shortened to 'Het' to not break formatting!
ApricotBananaGrape
Het Blue & IceHet Ice & BlueHet Denim & JadeHet Jade & Denim
Het Cucumber & SaffronHet Saffron & CucumberHet Intense Gold & EarthHet Earth & Intense GoldHet Intense Salt & MalachiteHet Malachite & Intense Salt
July
These eyes can only be found on July Celestial Lionesses and passed down through breeding.
DawnDraconidGalaxyNeutronPreonStarshineStellarSunglow
August
These eyes can only be found on August Maneater Studs or passed down through breeding.
Maneater

Skin Color
If you want to pass skins onto your cubs, I recommend using Rock Salts to guarantee all cubs in the litter having one specific skin color. These can be purchased in Monkey Business for 200

NCL Exclusive
These can only be found on NCLs, passed through breeding, or on lions born with a base tied to a specific skin.
Clouded Skin
Tied to: Ankh and MandarinPeach Skin
Tied to: No associated basesPale SkinTied to: Serruria and SphinxGreige Skin
Tied to: Medal, Murk, Nefer, and Phantom

Breed-Only Fail Skins
These skins can only be produced by parents with certain bases, or passed down through breeding. The bases required for each skin are listed below in bold, as well as the associated bases in plain text.
Echo Skin Interstellar
Tied to: NoneEggshell Blue SkinHaliotis
Tied to: JellyfishInk SkinNoctis or Styx
Tied to: noneLilac SkinGlass, Protea, or Kunzite
Tied to: nonePlum Skin Prismatic or Water Hyacinth
Tied to: NoneQuail Skin Buttercream, Pearl, Qahir, or Teardrop
Tied to: NoneSandstone Skin Arabica, Buff, Cameo, or Flaxen
Tied to: NoneWelsummer Skin Anjeer, Sapela, Sepia, or Shedua
Tied to: NoneBunting Skin Champagne, Citrine, Goldenrod, or Sulphur
Tied to: None

Event Skins
This skin can only be obtained by breeding to maneater studs in the August event or having a cub with the associated base.
Ruffian Skin
Tied to: Outlaw

Mane Shape & Mane Color
If you want to pass specific Mane Shape & Color to your cubs, your best bet is breeding two lions with the same Mane Shape & Color. Past that, you can use an Ochre Saltlick, which you can get from the September Prophet Shop, although it is not recommended for most breedings.

Mane Shapes
Combo Mane Shapes
These shapes can be bred by breeding certain factors together, or passed down through breeding! Factors are listed below in bold.
Royal ManeBarbary x Hellraiser, Sideburn, Thick, or Tufted
OR
Sideburn x Hellraiser, Thick, or TuftedBreed-Only Manes
These manes can be obtained through Maneater Studs in the August Event, or passed down through breeding!
Pariah ManeSavage ManeTsavo ManeMane Colors
All Mane Colors are shown on Regal mane, as that's what the wiki shows them on!
NCL Exclusive
These manes can be obtained through NCLs in Explore, or passed down through breeding!
Clay ManeCopal ManeMarigold ManeCombo Manes
These manes appear on 50% of all cubs born with the matching combo base, or can be passed down through breeding!
Cairngorm ManeCitrine ManeElysian ManeMadagascar ManeQahir ManeSerruria ManeSoul ManeBreed-Only Manes
These manes can only be passed down through breeding, and are usually spread via Raffle Lionesses.
Algae ManeAmber ManeAnjeer ManeAshen ManeBriar ManeCameo ManeChampagne ManeChartreux ManeCimmerian ManeCinnabar ManeCocoa ManeDinar ManeEbony ManeFiery ManeFlaxen ManeFlint ManeFulvous ManeGoldenrod ManeHoney ManeLilac ManeNacarat ManeOrchid ManePearl ManePistachio ManePlatinum ManePrune ManeSepia ManeShedua ManeSlate ManeTaupe ManeUmber Mane

Markings
While I have used images for everything else, there are literally hundreds of raffle markings and breed-only markings, so I will instead be using an image of the first marking of the shape group, and then listing all other colors beside them. You'll see what I mean soon! Markings will be shown on the base that I personally feel shows the contrast best, between Clear White or Styx.
This will also almost definitely be in multiple parts, so I apologize for the length.
PLEASE USE CTRL + F TO SEARCH VIA SHAPE NAME RATHER THAN FULL NAME
for example, if you were looking for red burmese, you would search burmese rather than 'red burmese'

Raffle Markings
Raffle Markings, aka Tier 2 Markings, can be found on Raffle Lionesses, May Groupies, potentially applied via RMAs, or passed down through breeding.
Shown: Briar Brindle
Other Colors: Bushveld, Fiery, Gold, Seal, Spice, & TangorShown: Black Burmese
Other Colors: Blue, Briar, Brown, Cimmerian, Dark Brown, Henna, Lilac, Noctis, Onyx, Red, Scoria, & SealShown: Algae Crackle
Other Colors: Almond, Blue, Briar, Brown, Bushveld, Cimmerian, Cinnabar, Dark Brown, Fiery, Goridhe, Lilac, Marigold, Noctis, Onyx, Red, Saffron, Seal, Spice, & WhiteShown: Algae Crumbing
Other Colors: Bushveld, Lilac, Pistachio, Tan, & TangorShown: Dim Black
Other Colors: Blue, Brown, Cimmerian, Dark Brown, Gold, Goridhe, Lilac, Mahogany, Noctis, Onyx, Red, Saffron, Seal, Silver, Spice, & WhiteShown: Black Dorsal Line
Other Colors: Blue, Briar, Brown, Bushveld, Cream, Dark Brown, Gold, Lilac, Noctis, Onyx, Red, Seal, Spice, Tangor, & WhiteShown: Feline 1 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 2 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 3 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 4 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 5 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 6 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 7 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 8 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Feline 9 Black
Other Colors: Dark Brown, Fiery, Ginger, Gold, Lilac, Noctis, Onyx, Silky, & SilverShown: Briar Freckles 1
Other Colors: Bushveld, Ginger, Lilac, Seal, & SpiceShown: Briar Freckles 2
Other Colors: Bushveld, Ginger, Lilac, Seal, & SpiceShown: Briar Freckles 3
Other Colors: Bushveld, Ginger, Lilac, Seal, & SpiceShown: Fulvous Glaze
Other Colors: There are currently no other BO Glaze Raffle Markings!Shown: Almond Indri
Other Colors: Blue, Bushveld, Chert, Clay, Cream, Dark Brown, Ginger, Fiery, Gold, Goridhe, Lilac, Noctis, Onyx, Red, Saffron, Scoria, Seal, Steele, & WhiteShown: Almond Inverted Zebra
Other Colors: Bone, Cream, Marigold, Pistachio, Quartz, Royal, Saffron, Shell, & WhiteShown: Almond Lace
Other Colors: Bone, Coral, Cream, Marigold, Pistachio, Quartz, Royal, Saffron, Shell, Silky & WhiteShown: Leg Banding Auburn
Other Colors: LilacShown: Algae Margay
Other Colors: Blue, Briar, Bushveld, Chert, Cimmerian, Cinnabar, Cream, Dark Brown, Fiery, Ginger, Goridhe, Lilac, Noctis, Onyx, Red, Saffron, Seal, & SteeleShown: Algae Panther
Other Colors: Black, Briar, Bushveld, Chert, Cimmerian, Clay, Dark Brown, Gold, Noctis, Onyx, Red, Scoria, Seal, Spice, Steele, & TangorShown: Algae Pelage
Other Colors: Almond, Black, Brown, Bushveld, Chert, Cimmerian, Dark Brown, Fulvous, Goridhe, Lilac, Noctis, Onyx, Saffron, Seal, Spice, & SteeleShown: Lilac Points
Other Colors: There are currently no other BO Points Raffle Markings!Shown: Bushveld Quail Flecks
Other Colors: Goridhe & LilacShown: Almond Rims
Other Colors: Bone, Cream, Marigold, Pistachio, Quartz, Royal, Saffron, Shell, Silky, & WhiteShown: Auburn Roan
Other Colors: Bushveld, Fiery, Ginger, Goridhe, LilacShown: Algae Rogue
Other Colors: Black, Bushveld, Chert, Cimmerian, Dark Brown, Fulvous, Goridhe, Lilac, Noctis, Saffron, Scoria, Seal, Spice, Steele, & TangorShown: Fulvous Shine
Other Colors: PistachioShown: Bushveld Siamese
Other Colors: Goridhe, Lilac, & Seal

Markings Continued
Raffle Markings
Shown: Almond Soft Unders
Other Colors: Bone, Coral, Cream, Marigold, Pistachio, Quartz, Royal, Saffron, Shell, Silky, & WhiteShown: Tail Banding Auburn
Other Colors: LilacShown: Almond Underfelt
Other Colors: Bone, Coral, Cream, Marigold, Pistachio, Quartz, Royal, Saffron, Shell, Silky, & White
Vitiligo 1Vitiligo 2Vitiligo 3Vitiligo 4Vitiligo 5Vitiligo 6Vitiligo 7Vitiligo 8Vitiligo 9Rosettes
These markings can appear randomly on cubs or be passed down through breeding! They can also very rarely be gotten via RMAs.

Shown: Auburn Heavy Rosette
Other Colors: Briar, Bushveld, Dark Brown, Doubloon, Ebony, Ginger, Goridhe, Noctis, Onyx, Red & Seal
Shown: Inverted Heavy Rosette Bone
Other Colors: Cream, Fiery, Gold, Saffron, Silky & White
Shown: Inverted Rosette Cream
Other Colors: Fallow, Fiery, Gold, Saffron, Silky & White
Shown: Bone Inverted Shaded Rosette
Other Colors: Cream, Fiery, Gold, Saffron, Silky & White
Shown: Bone Inverted Shaded Rosette
Other Colors: Cream, Fiery, Gold, Saffron, Silky & White
Shown: Briar Rosette
Other Colors: Brown, Bushveld, Cream, Dark Brown, Fiery, Ginger, Gold, Goridhe, Noctis, Onyx, & Seal
Shown: Auburn Shaded Rosette
Other Colors: Briar, Bushveld, Dark Brown, Doubloon, Ebony, Fiery, Ginger, Goridhe, Noctis, Onyx, Red, & Seal
Shown: Auburn Soft Rosette
Other Colors: Briar, Bushveld, Dark Brown, Doubloon, Ebony, Fiery, Ginger, Goridhe, Noctis, Onyx, Red, & Seal

Hybrid Markings
These markings can only be found on First Generation Leopons or Tigons and their descendants!
Mottled Rosette
Leopon ExclusiveMottled Stripes
Tigon Exclusive

Markings Continued
NCL Exclusive Markings
These markings can be found on NCLs, or be passed down through breeding.

Shown: Clay Back
Other Colors: Copal
Shown: Cinnabar Belly
Other Colors: Marigold
Shown: Clay Brindle
Other Colors: Copal
Shown: Cinnabar Coat
Other Colors: Copal
Shown: Cinnabar Face
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Inverted Brindle
Other Colors: Marigold
Shown: Almond Inverted Cover
Other Colors: Bone, Cinnabar, Coral, Cream, Marigold, Quartz, Royal, Saffron, Shell, Silky, & White
Shown: Leg Banding Cinnabar
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Luster
Other Colors: Marigold
Shown: Cinnabar Marozi
Other Colors: Copal
Shown: Clay Mask
Other Colors: Copal
Shown: Clay Paws
Other Colors: No other colors
Shown: Cinnabar Points
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Roan
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Sable
Other Colors: Clay, Copal, & Marigold
Shown: Clay Shell
Other Colors: No other colors
Shown: Clay Shepherd
Other Colors: No other colors
Shown: Cinnabar Smudge
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Smudge
Other Colors: Clay & Copal
Shown: Cinnabar Socks
Other Colors: Marigold
Shown: Tail Banding Cinnabar
Other Colors: Clay, Copal, & Marigold
Shown: Cinnabar Underbelly
Other Colors: Marigold
Shown: Cinnabar Undercoat
Other Colors: Marigold
Shown: Cinnabar Unders
Other Colors: Marigold
Shown: Cinnabar Undersides
Other Colors: Marigold
Shown: Cinnabar Undertone
Other Colors: Marigold
Shown: Cinnabar Vesture
Other Colors: Marigold

Markings Continued
Event Markings
These can be found through mechanics in their respective events, namely, July Celestial Raffle Lionesses or August Maneater Studs!

July
Celestial AgoutiCelestial CoverCelestial ShineCelestial SpecklesDawn DappleDawn LaceDawn MargayDawn ShineLeonid CoatLeonid FelineLeonid SmokeLeonid UndershinePrismatic FelinePrismatic GlazePrismatic SmokePrismatic UndercoverSidereal CoverSidereal GlazeSidereal PointsSidereal ShineSkyward CoverSkyward CrumbingSkyward ShineSkyward UndercoverStratosphere FelineStratosphere LycaonStratosphere UndertoneStratosphere Vitiligo

August
Bandit CloudBandit FelineBandit MirageBandit UndersOutlaw CheetahOutlaw FelineOutlaw Mottled FissuresOutlaw RogueRuffian FelineRuffian PantherRuffian SiameseRuffian Soft UndersScoundrel FelineScoundrel Inverted BrawlScoundrel PelageScoundrel UndersidesUkame FeralisUkame MarbleUkame Rugged UndersUkame UndercoverVagabond MirageVagabond Mottled FissuresVagabond NuzzleVagabond Reverse Indri

Year-Round Bases
These bases can be bred for year round, and are not limited in any way. Monthly Combos are in the Event Base section. Base Genetics are listed under each base! S stands for Solid, and CS stands for Countershaded

Rare Bases
These bases are only available as fails from any breeding involving an Uncommon or higher rarity.
Amber
Cream Dark SAnjeer
Red Medium CSArgent
Cream Medium SAshen
Black Light SBuff
Cream Light CSButtercream
Cream Medium SCameo
Cream Medium CSChampagne
Cream Light CSChartreux
Black Light CSCinnabar
Red Light SCocoa
Black Medium CSDinar
Gold Dark CSEbony
Black Dark SFiery
Red Light CSFlaxen
Cream Dark CSFlint
Black Medium SFulvous
Gold Dark CSGoldenrod
Gold Medium SGoridhe
Gold Medium CSHoney
Gold Dark SMaroon
Red Light CSNacarat
Gold Medium CSNoctis
Black Dark SPlatinum
Black Medium SPrune
Red Dark CSSapela
Red Light SShedua
Red Dark SSlate
Black Dark CSSulphur
Gold Light STaupe
Red Light STeardrop
Gold Light SUdara
Gold Dark SUmber
Red Medium S
Raffle Bases
These bases can only be obtained via Raffle Lionesses and are passed down through their offspring.
Bushveld
Gold Medium CSHaze
Black Dark SLilac
Black Medium SOrchid
Red Light SPearl
Cream Medium SPecora
Cream Medium CSSepia
Red Medium S

Hybrid Bases
These bases can only be found on First Generation Leopons or Tigons and their descendants!
Kimanjano
Leopon ExclusiveMandarin
Tigon Exclusive

Event Bases
These bases can be obtained in their respective events, and passed down through offspring that way!

May
These bases are obtained from May Groupies
Abyssinian
Gold Dark CSDamu
Gold Dark SNudar
Gold Light CSRipe
Gold Dark CSSahara
Gold Medium SSunspot
Gold Medium CSWine
Red Medium CS
July
Celestial
Black Dark CSDawn
Red Light CSJellyfish
Black Dark CSLeonid
Black Medium CSPrismatic
Red Dark SSidereal
Black Medium CSSkyward
Black Light CSStratosphere
Black Light CSSunrise
Red Light CSSunset
Red Medium CS
August
Bandit
Red Light CSOutlaw
Cream Dark CSRuffian
Cream Medium SScoundrel
Red Medium CSUkame
Gold Light CSVagabond
Cream Dark CS

Combo Bases
Due to size limitations, I will not be adding their factors. Instead, you can visit the Combo Base wiki page in order to see the factors of all Combo Bases.

Time Limited
These bases can only be bred during their respective months.
Mistletoe - Dec
Black Light CSPulsar - July
Black Light CSTemporal - Oct
Red Dark CSWitch - Oct
Black Dark CS
Seasonal or Time of Day
The required time of day and/or season will be in the name! means Dry Season, means Wet Season, means Day Time, and means Night Time. Please note that Day/Night goes off of LDT, not your personal timezone.
Cloudburst
Black Medium CSEsker
Cream Medium CSHoarfrost +
Black Medium CSMajivu
Black Medium SRime +
Black Medium CSSerruria
Red Light SSmog
Cream Medium CSSnowflake
Black Light CS
Standard Combo Bases
Arabica
Cream Dark CSAsiatic
Cream Medium CSBasalt
Black Medium SBlack Rose
Red Dark SCairngorm
Black Light SCelsian
Black Light SCitrine
Gold Light CSElysian
Black Dark CSEmber
Gold Dark CSEnnead
Black Medium CSGilded
Gold Medium SHellebore
Cream Dark SHexaplex
Gold Light CSMadagascar
Red Dark CSMudstone
Gold Dark SNadir
Black Dark SOgdoad
Red Medium CSOlive
Cream Dark CSQahir
Cream Dark SRa
Black Dark CSRose Gold
Red Medium SSoul
Black Light CSSphinx
Cream Medium SSquall
Red Medium CSStyx
Black Dark S