It really depends! Different people look for different things, if youre looking to stud out this heir, then markings seem to work for me as I get a ton of stud requests (Although my queen in question is 9xbo and a special base ^^) But if youre not gonna stud out this king, then choose which one you like more, I find that smilus doesn't sell too well unless it has something else to it. So your best bet is to go with the special base or markings if you want the best profit. It also depends on if the heir in question is pleasing to look at, i've found lions with tons of bo but i'd never stud to them because they look muddy and i'm thinking about the cubs that would be produced.

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If you’re looking to stud this guy out it’d say base is the first factor people look for - combo and breed only
Then markings - they will help you stand out from the other studs with the base
Mutation is a bonus But a good one, pies are the usual - if you want to go primal - ferus is good since people will search for a ferus king for ferus laciná due to pennyroyal effect :))

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Id say a Special Base, especially a combo factor one, (Ice has many but this also means there might be quite a few Ice Base studs ofc) and BO Traits for the Base & Markings
For markings, you could also include a rosette or two! (People want (non-inverted) rosettes, you have rosettes, you just might make some money)
As for mutation, I usually see Pies/Patches paired with Rosettes! (Plus it looks pretty damn good most of the time!) If your looking for sumth that isnt Patches/Pies, I would try a ferus! (Due to Pennyroyal Chances! Ferus's also have some bonuses unlike other Primal Muts (Eg Small things like Extra Damage against bipedial creatures, ability to pass genes from both genders)
~~remember to make it look good too! (No clashing markings etc etc)
~~people will usually (I think-) go for clean lions over dirty ones!

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