The tutorial has gone unchanged since it's creation, back when lioden was a very different game functionally! I feel a lot of improvements and updates could be made to help new players get to grips with things.

Replayable Tutorial
For both new and returning players, the tutorial may be a lot to get to grips with in one sitting. The fact it can't be replayed puts players who have gone on hiatus at a dis-advantage and it can lead to some players never returning because they can't get the hang of things.

Episodic/Quest style tutorial
Sitting through a whole tutorial from start to end can be a strain, so re-playable sections in the form of quests or episodes would be a good way to refresh on specific subjects.

Full tour of the site
A number of newer features are left out of the tutorial, including the adventure page, search function, gorilla enclave and giving tree. There is also no explanation of the different options for the chat box, leading to few new players knowing about sales chat to advertise things in the correct place or the help box to seek support.
Strictly requiring players to visit each broad section of chatter as well as other locations mentioned but not visited in the tour would be beneficial also

Genetics
A general run down of genetics including marking tiers, base rarity meanings and how mutations work wouldnt go amiss. Even providing a direct link to their sections on the wiki would allow those who need to know more about genetics to learn before starting.

More active participation tasks
A lot of the tutorial waters down to "go to place, read about place" without prompting interraction with features. Other similar sites prompt new players to make a comment on an "introduce yourself" forum, run through a quest for an npc (ie gorilla enclave) or join a clan to confirm understanding of each feature.

Broodmothers
Broodmothers are not covered whatsoever in the tutorial, leaving new players oblivious to their function of keeping cubs alive. This could easily be merged with the breeding tutorial- go claim another lioness. She has a nurturing personality and would love to babysit your cubs. Set lioness to broodmother and assign cub to lioness, Nice and safe!

Territory
The shifting mechanic for territory growth is a confusing one in general, a brief pop up tutoral would be helpful to explain the mechanic to players as and when it changes from paid, to pvp and back to paid slots again.

TLDR:
Replayable tutorial as episodes or quests
Cover all the areas of crossroads.
Explanation of chat sections and brief rundown of rules
Explanation of genetic traits
Active participation in tutorial instead of reading.
Broodmother tutorial
Territory Tutorial.

I'd love to hear feedback on this suggestion! If you support or not I'd love to know why, but remember to keep comments respectful to both new and experienced players

More suggestions
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Dedicated Help Chat Helpers
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Tutorial Re-Haul
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Personality Suggestion- Impatient
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Personality Suggestion- Diligent
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Un-pin abandoned posts
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An option to opt-out of post-hiatus needs refills