I tried to find some info and I'm really lost on what applicators can be passed onto cubs and what not, or is is that none can be passed down.

In fact, what is considered an applicator and what not? Does the name need to be included in the item, such as "Applicator: Hyena Stripes Heavy"? Are those that don't have the name "applicator" inheritable, such as Astral Blessing, Augur Fur, Brimstone Imprints, Penumbra Body or Frostbite Protection?

These are some of the items I found in my hoard under the category "Applicator", but I don't know which of these my cubs can inherit from their parents and what not. What is the easiest way to figure out what applicators can be passed down, or does it literally vary from item to item? And for that matter, if I find a female that has markings that were gained through applicators, how do I know these can be passed to cubs?

I'd hate to waste these on my future king, or keep certain females, just to later learn that their markings or colors can't be passed onto their children.